Morgan observed tight linkage in Drosophila with only 1.3 percent recombination for

1.  White eye and yellow body

2.  Normal wing and yellow body

3.  White eye and miniature wing

4.  Yellow body and miniature wing 

Morgan coined the term recombination to describe 

1.  Physical association of genes on a chromosome

2.  Generation of new traits by mutation

3.  Linkage of two genes on different chromosomes

4.  Generation of non-parental gene combinations